ニホンヒキガエルBufo japonicus japonicusの自然誌的研究 : IX.繁殖期における♂の行動

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The reproductive behavior of the male Japanese toad, Bufo japonicus japonicus, was observed at several ponds in Kanazawa prefecture, central Japan. In the two ponds where males outnumbered females by six to one, 88% of the male toads stayed on the land around the pond, waiting and searching for females. On the other hand, in another pond, where males and females were nearly equal in numbers, 44% of the males waited and searched for females in the pond. The males' waiting position on land was located on a line between their place of residence and the pond. Males stayed longer in the breeding pond (5 to 6 days) than females (1 to 2 days). The maximum record of breeding times was 7 times in 8 seasons for a male, and 4 times in 6 seasons for a female.
1985-03-30