ミズゴマツボ科の類縁関係

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The stenothyrid group has been classified in several different family-subfamily group according to the view point of the criteria which are adopted by different authors. There has been a considerable difference of opinion concerning the proper family and subfamily designation for Stenothyra since 1856. Most authors have treated stenothriid snails as animals closely related to the Hydrobiidae, Bithyniidae or Rissoidae. The general anatomy of the European Bithynia tentaculata was presented by Lilly (1953) where she described precisely the radula and genital organs beside other ecological and anatomical features. The features of Stenothyra are rather different from those of Bithynia, e. g. the rachidian of the radula of the latter has no basal cusps and the lateral tooth also differs in general outline ; genital organs are different in several points such as the presence of a prostate organ and the absence of the female gonopericardial duct; the operculum of the latter has no elevated ridge on the inner surface and is calcareous and concentric type. Bithynia also differs in the appendage of verge with flagellum, no spermathecal duct etc. Therefore it seems that there is no close relationship between Stenothyra and Bithynia. In Alvania as well as in the Hydrobiidae the pallial vas deferences is developed into a prostate, in Stenothyra. Rissoa and Rissoina on the other hand, with the exception of R. inconspicua, this structure is non glandular and the penial duct which consists of a secreting epithelium functions as a prostate. These circumstances seems to confirm the opinion that the Stenothyriidae is closely related to Rissoa and Rissoina than to the Hydrobiidae : on the other hand it means that the hydrobiid snails are more closely related to Alvania than to other rissoids ridges which found commonly through Rissoa and Rissoina. The rachidian tooth of Stenothyra has basal cusps on the anterior surface thus distinguishing it from rissoid snails where the basal cusp is situated rather posteriorly. The female genital organs are rather similar in these three groups. However, the upper oviduct gland which is found in European Alvania and Rissoa can not detected in Stenothyra. nd Hydrobia. The lack of an upper oviduct gland is perhaps the most conspicuous feature which distinguishes the Hydrobiidae from the Rissoidae. In the view points of the genital organs, radulae, operculum and other anatomical features, the Stenothyridae has a close relationship with both the Rissoidae and Hydrobiidae and possibly originated from the common ancestral stock.
日本動物分類学会の論文
1969-10-01

ミズゴマツボ科の類縁関係

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