NAKAOKA Yuji | Hokkaido Nemuro Livestock Hygiene Service Center, 69 Betsukaimidorimachi, Betsukai, Notsukegun, Hokkaido 086–0214, Japan

Asahikawa Asahiyama Zoological Park and Wildlife Conservation Center

Asahikawa Asahiyama Zoological Park And Wildlife Conservation Center

Department Of Animal Science And Resources Science College Of Bioresource Sciences Nihon University

Asahiyama Zoological Garden

Department of Wildlife Science, Nihon University, 1866 Kameino, Fujisawa, Kanagawa 252–8510, Japan

National Institute of Infectious Diseases, 1–23–1 Toyama, Shinjuku–ku, Tokyo 162–8640, Japan

Hokkaido Kamikawa Livestock Hygiene Service Center, 4–15 Higashitakasu, Asahikawa, Hokkaido 071–8154, Japan

Hokkaido Nemuro Livestock Hygiene Service Center, 69 Betsukaimidorimachi, Betsukai, Notsukegun, Hokkaido 086–0214, Japan

Asahikawa Municipal Asahiyama Zoological Park & Wildlife Conservation Center, Kuranauma, Higashiasahikawa-cho, Asahikawa, Hokkaido 078–8205, Japan

Asahikawa Municipal Asahiyama Zoological Park & Wildlife Conservation Center, Asahikawa 078-8205, Japan