ヤマゴボウ属の花の構造およびナデシコ科の花との関連
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概要
- 論文の詳細を見る
In the genus Phytolacca, the flower of P. dioica seems to have the most primitive structure, though it is specialized in being unisexual. This male flower consists of 5 perianths, 20-30 stamens and 3-5 sterile carpels. The stamens are generally arranged in 10 fascicles of which the five fascicles (dst) situated on the dorsal side of the carpels with 1-2 stamens each and the five fascicles (1st) on the lateral side of the carpels with 3-4 stamens each (Figs. 1 and 3:4). One of the observed flowers has 29 stamens which are grouped into 12 staminal fascicles (Fig. 2:2 and 3:3). They are five fascicles of (dst), five fascicles of (1st) and two additional fascicles of (ast) (Fig. 3:3). Since the female flowers have an 8-10-carpellate ovary, it is possible that the male flowers having an 8-10-carpellate ovary and 16-20 staminal fascicles are assumed to be the basic structure. The normal male flower seems to be derived from the basic structure by reduction of staminal fascicles into 10. The two additional fascicles of (ast) seem to be the remains of the staminal fascicles of the assumed basic male flower. In the other flower observed (Fig. 2:1), it has 23 stamens in which each of two fascicles of (dst) consist of 3 stamens as compared with one or two stamens in the normal flower. These facts indicate that the basic structure of the flower in the genus Phytolacca is assumed to have 5 sepals, 16-20 fascicles with 3-4 stamens each and an 8-10-carpellate ovary as seen in Fig. 3:5. In P. japonica the normal flowers have 5 perianths, 7-9 stamens and a 7-9-carpellate ovary. The stamens are usually situated on alternate with the carpels (Fig 4:1). One of the observed flower has 5 perianths, 11 stamens and an 8-carpellate ovary (Fig. 4:2). Among these 11 stamens, the eight are situated on the usual position of alternate with the carpel, two are derived from the division of each of two normal stamens, the remained one (dst) is opposed to the carpel. The other flower has 6 perianths, 13 stamens and a 7-carpellate ovary (Fig. 5). among these 13 stamens, the seven (1st) are situated on usual position of alternate with the carpel, the five (dst) are opposed to the carpel and the remained one is divided from the basic structure of the genus by reduction of the number of stamens in the staminal fascicles of (1st) and entire disappearance of the staminal fascicles of (dst).
- 日本植物分類学会の論文
- 1987-09-25